Making this a “snapshot” is my way of dealing with the overwhelming diversity of butterfly species one encounters over two weeks and across 10,000 vertical feet in the cloudforest on the Amazon side of the Andes Mountains. During this terrific adventure, I photographed 35 hairstreak species, 65 metalmark species, about 250 different brushfoots species, including 90 satyrs, about 20 pierids and swallowtails, and about 100 different species of skippers. Getting there and taking the photos was relatively easy. Then came the sorting of more than 4,000 images and identifying what was photographed, a task which will never be complete. A trip lasting 2.5 weeks basically swallowed up 4 months of my mental capacity for butterflies.
This trip was organized by Mike McInnis and co-led by Peru’s own Dr. Gerardo Lamas. Other participants were Dr. Olaf Mielke, Dr. J. D. Turner, Steve Kinyon and Bill Dempwolf, all shining lights in the butterfly world. This was year 17 (I think) of the project to identify all the butterflies in the Cosñipata Valley study area, on the edge of Manu Parque Nacional. For some geographic scale, Manu National Park is about the size of Socorro County, NM. The area we traveled in was perhaps 50 miles in length and spanned about 9000 feet of elevation, comparable to traveling from the summit of New Mexico’s Sierra Blanca downstream to Roswell. Their working species total was – are you sitting down – more than 2600 species. All other participants were collecting butterflies whereas I was mostly taking photos.
A second thing to keep in mind is that virtually all our interactions with butterflies were enabled or facilitated by the team’s use of various baits or traps. Spitwads were favored by skipper lovers. The rest of us, every day, fabricated and sprayed onto trailside vegetation a liquid mixture of urine (electrolytes) and rotting fish (the odor that draws them in). Such practice is not typical in the US; I never use baits here in NM because most butterflies come readily to flowers at eye level or below. But in South America, simply walking and observing in unbaited habitat might get you 10% of all the species that are there. Most are up in the trees and thus inaccessible to earth-bound humans. Many collectors use long-handled nets. Bait provides highly prized electrolytes which draw the vast majority of butterflies down to earth and holds them still while you watch, click, or swing.
I organized the presentation below in a rather uncomplicated way mostly by offering photos with captions, in the chronology in which they were taken. I think most of you want to see the images, so I focused (so to speak) on that, but I also try to offer informative captions. After I returned home, I received Volume I of David Geale’s highly recommended “Amateur’s Guide to the Butterflies of Eastern Ecuador & Peru: Papilionidae, Pieridae, Nymphalidae” hot off the press, so I use his English names when offered [Volume II will have the Hairstreaks, Metalmarks and Skippers]. The identifications are mostly mine, but with generous help on difficult species from the entire team. The misidentifications are all mine, but I hope few in number. Please correct me if you see an error!
Cherry Saqra (Pedaliodes tucca: Nymphalidae: Satyrinae) 1900m along Manu Road; October 9, 2024 (photo by Steve Cary). Our descent from Cusco into the Cosñipata valley started high (like most descents). Weather was poor at the highest potential stops, so we missed a chance for high elevation hairstreaks in the Penaincisalia genus. Continued descent was accompanied by warming temperatures. Though still cloudy when we stopped at 6200′ to wait for a road maintenance crew to let us by, we exited the van and saw a handful of things. At that elevation, satyrs are numerous. I photographed two including the above.
You may scratch your head and wonder how a place as small as Peru could possibly have 5,000 butterfly species. Well, the genus Pedaliodes, has 176 species already described, with 92 new species in the process of being described. When you digest that, go ahead and ask me about related genera including Altopedaliodes, Antopedaliodes, Corderopedaliodes, Neopedaliodes, Panyapedaliodes, Parapedaliodes, Pherepedaliodes, Physcopedaliodes, Protopedaliodes, Punapedaliodes and Steromapedaliodes.
Our first full day of field work was October 10, and we naturally went to the place people had been talking about for the last day and a half – Quebrada Quitacalzon. Quebrada is the local term for small, steep drainages tributary to larger watercourses. Our road down from Cusco crossed many of these. They are good places to look for butterflies because they often have parking pullouts, bridge abutments, and especially damp soils and creek banks. Quebrada Quitacalzon is the most famous of these in our group because – fasten your seatbelts – they have documented more than 1,000 butterfly species there over the 17 years of study. The entire walkable area has to be less than 1 square mile, but there are several creek crossings and a short (100-yard) trail leading into the forest. Nevertheless, that is remarkable species richness for such a small area.
Epione Sister or White-Banded Sister (Adelpha epione agilla: Nymphalidae: Limenitidinae) Quebrada Quitacalzon, October 10, 2024 (photo by Steve Cary). This was perhaps the most distinctive of the ~15 different species of sisters (Adelpha sp.) that I photographed personally. One of 44 different sister species documented from the Cosñipata Valley. One of perhaps 250 species known from the Western Hemisphere.Two-Barred Flasher complex (Telegonus fulgerator complex: Hesperiidae: Eudaminae) Quitacalzon; October 10, 2024 (photo by Steve Cary)Copper-headed Sootywing (Bolla cupreiceps: Hesperiidae: Pyrginae) Quebrada Quitacalzon; October 10, 2024 (photo by Steve Cary). The study area was home to a large handful of various sootywing species, all very similar to our own Staphylus species. This one was the most distinctive. Reminded me of New Mexico’s Staphylus ceos.Magenta-banded Leafwing (Fountainea nessus: Nymphalidae: Charaxinae) Quebrada Quitacalzon; October 10, 2024 (Photo by Steve Cary). It was thrilling to watch this individual perch and bask laterally for a couple minutes, then gradually turn to align itself with the sun, then slowly open its wings to reveal an iridescent magenta/blue sheen that must really get the attention of nearby females. Quite a show!no English name (Caeruleuptychia tenera: Nymphalidae: Satyrinae) Quebrada Quitacalzon; October 10, 2024 (photo by Steve Cary). As best I can ascertain, the genus Caeruleuptychia was new for me despite three prior trips to Ecuador and Peru. Our leader, Dr. Gerardo Lamas, seemed particularly fond of this genus, which seems endemic to South America, and I appreciated the extra effort he made to point out details of wing marks. We might think that all the blue satyrs we saw were the same species, but in fact the underside ocelli vary quite a bit. I think I photographed six different species, and this was my favorite.Ornate Calligraph or Godart’s Numberwing or Pygas Eighty-eight (Catagramma pygas: Nymphalidae:Biblidinae) Quebrada Quitacalzon; October 10, 2024 (photo by Steve Cary)Frosted Giant Owl (Caligo idomeneus: Nymphalidae: Morphinae) Quebrada Quitacalzon; October 10, 2024 (photo by Steve Cary). Okay, not the greatest photo ever of a hand-sized “owl butterfly.” I was pleased with myself for spotting this guy in the roadside underbrush, then patiently waiting while it fidgeted around, probably trying to access my spilled bait. I snapped a sharp broadside view (not shown here, sorry) then finally it opened its wings enough for what I consider to be a rare view of the upperside, which simplified the identification to species.
I photographed about 135 species at Quebrada Quitacalzon that day. The next day we all went farther up the mountain, while splitting up a bit. I went with the van to a place called Yanomayo, which was familiar to veterans of this study, perhaps 7500 to 8000 feet elevation.
Cupavia Skipperling (Dalla cupavia: Hesperiidae: Heteropterinae) Yanomayo; October 11, 2024 (photo by Steve Cary). This location, at least near wet spots, was frequented by a few species of Dalla, which are in the skipperling subfamily along with our New Mexico friends Piruna pirus and Piruna polingii.Darkened Rainbow-Saqra or Rusty Mountain Satyr (Lymanopoda ferruginosa Nymphalidae: Satyrinae) Yanomayo; October 11, 2024 (photo by Steve Cary). Notice the red mite on a wing vein. This higher country was rich with butterflies, but with different groups and species than you might imagine. In addition to the skipperlings like Dalla spp., above, there was an abundance of satyrs. I show you this one so I can rave about the species richness of the Lymanopoda genus: 60 described species with 5 new species in the works.
The next day was an opportunity to head back down the hill to our lowest location yet, the Chontachaka Ecological Reserve, at about 2500 feet elevation, I think it was completely new spot for the group. Dr. J. D. Turner, our metalmark guru, had been investigating the place on his own for a few days, studying his “Rios.” He thought it might be a good place for our whole group and he was right!
Godman’s Sarota (Sarota myrtea: Riodinidae: Riodininae) Chontachaka; October 12, 2023 (photo by Steve Cary). The metalmarks indeed were glorious, if tiny. Of the 65 species I photographed, this was the most exciting one for me because I had botched previous attempts years ago to photograph something in this genus: hairy legs and scintillating silver scales. This one plopped down in front of me, right on the trail edge, no bait involved. My first shot was from the rear (pretty embarrassing), but then it turned to bask laterally, offering a side view. I clicked off three more flash-assisted shots and was a happy camper.Olive Metron (Metron zimra: Hesperiidae: Hesperiinae)Rust-Coloured Round-Owl (Catoblepia berecynthia adjecta: Nymphalidae: Morphinae) Chontachaka; October 12, 2024 (photo by Steve Cary).Celmus Hairstreak (Celmia celmus: Lycaenidae: Theclinae) Chontachaka; October 12, 2024 (photo by Steve Cary) This may be the most common and widespread hairstreak in Amazon Basin. I include the image because it represents a landmark moment for me. For the first two field days, I struggled to even see the tiny hairstreaks, let alone photograph them. In my defense, there were a lot of flashy colors dodging about (see photos above) and it takes a lot of mental discipline not to follow the neon signs with your eyes.Sista Hairstreak (Janthecla sista: Lycaenidae: Theclinae) Chontachaka; October 12, 2024 (photo by Steve Cary). Finally, I managed to focus on the tiny hairstreaks that also were visiting baited leaves. They twinkled silver and blue (males anyway) when arriving and leaving, but while on the baited leaf they did not draw attention to themselves. I was very relieved to figure it out!
Rocotal is a terrific butterfly spot that is about halfway up the mountain at about 6000 feet elevation. It has a large pullout, a rain shelter, and multiple creek crossings if you are willing to walk a couple miles along the road. Rocotal was part of our deployment on October 13, and that’s where I spent most of my time.
Red-Spotted Satin-Wing or Leprieur’s Glory (Asterope leprieuri: Nymphalidae: Biblidinae) Rocotal; October 13, 2024 (photo by Steve Cary).Carmine Falcon-Saqra (Corades enyo: Nymphalidae: Satyrinae) Rocotal; October 13, 2024 (photo by Steve Cary). Just as with satyrs in New Mexico and the US, they give you undersides all day, but you have to be patient and lucky to get a good upperside view or photograph.Celestial Perigraph (Perisama philinus: Nymphalidae: Biblidinae) Rocotal; October 13, 2024 (photo by Steve Cary).Flame-Based Perigraph (Perisama calamis (Nymphalidae: Biblidinae) Rocotal; October 13, 2024 (photo by Steve Cary). Oh boy, another source of electrolytes!Foothill Jadewing or Green Longwing (Philaethria ostara Nymphalidae: Heliconiinae) Rocotal; October 13, 2024 (photo by Steve Cary). Kind of shaped like a swallowtail, but it’s a longwing; kind of resembles the also very emerald green Malachite, but it is its own beautiful thing.Undescribed Ruby-Eye (Perichares n. sp. Hesperiidae: Hesperiinae) Rocotal; October 13, 2024 (photo by Steve Cary). One thrill of the trip was this lovely skipper. Skipper lovers Bill and Olaf got busy walking the roadsides and putting out their spit wads before the sun even burned off the morning clouds. I was following and putting out fish bait. This may have been my first butterfly of the day, which I photographed as it investigated Bill’s salty offering. My flash showed why these skippers are called ruby-eyes. I snapped only two shots and was satisfied with the results. Bill was not far behind with his net. That evening Bill showed Olaf the specimens and Olaf promptly declared it was a completely new species. Olaf would know. I was pleased that Bill took the specimen because you cannot do complicated taxonomy on a photo, whereas a specimen allows for lab work to be done as needed.
The next phase of our trip took us to our lowest elevation location for a five-day stay. Pantiacolla Lodge was accessible by boat along the Rio Madre de Dios. To get there we packed up our gear, climbed into the van and drove downhill to the small town of Pillcopata, where we purchased some necessities. From there it was a short drive to the river and the businesses that served traffic on the river. Our destination, Pantiacolla Lodge, ran its own boat, so we piled in us and our stuff and motored comfortably downstream. The river we navigated is a tributary to a tributary to a tributary of the Amazon (or something like that). We were in the Amazon drainage network, which we had been all along of course. Pantiacolla is at about 1350 feet elevation, but still officially in the mellow cloud forest. Farther downstream is the lowland jungle, which has more snakes, disease-carrying insects, etc.
unknown clearwing (Nymphalidae: Ithomiinae) I largely ignored the clearwings, a hugely significant group of tropical butterflies that one cannot see anywhere else. They are difficult to identify and I simply had other priorities. But you need to see one and here it is.
Cleopatra Shadeflitter (Taygetis cleopatra male (left) and female (right): Nymphalidae: Satyrinae) Pantiacolla; October 14, 2024 (photos by Steve Cary). I thought that the number and variety of satyrs would decline as we went to lower elevations. Boy, was I wrong! There were many new genera and many lovely, large brown things flopping through forest. Here are two that I first thought were separate species, but they both keyed to Cleopatra; Gerardo confirmed they were male and female of this sexually dimorphic species.
Southern Amber-Phantom or Yellowish Clearwing-Satyr (Haetera piera: Nymphalidae: Satyrinae) Pantiacolla. This was my first of the Ghost Satyrs or Phantom Satyrs which are largely transparent and dance along forest trails.Shining Purplewing or Alpais Purplewing (Eunica alpais: Nymphalidae: Biblidinae) Pantiacolla; October 15, 2024 (photo by Steve Cary). I admit it is not an ideal photo background. At Pantiacolla our cabins were built safely off the ground and netted to keep out undesirable insects, which they did with 99% effectiveness. But doors do get opened and our cabins worked a little bit as large traps for the occasional butterfly. Bill and I found this inside our cabin when we returned from a day on the trails. It was a new species for me and the best view I’ve ever had of an upperside of any of the purplewings (Eunica species). Shining indeed!Red Commoner or Red Banner (Callicorina pulchra pallidior: Nymphalidae: Biblidinae) Pantiacolla; October 15, 2024 (photo by Steve Cary). The name is quite an understatement considering the radiant blue/pink/red that shimmer off its dorsumRainbow Agrias or Claudina Agrias (Agrias [Prepona] claudina: Nymphalidae: Charaxinae) Pantiacolla; October 16, 2024 (photo by Steve Cary). This butterfly was a semi-holy grail species because of its bright red and blue upperside combined with its assertive flight and its propensity to land/perch too high up on tree trunks to allow for decent photography. A day earlier I had a major run-in with chiggers on the grounds of our lodge. Not wanting a repeat when I went into the field this day, I impregnated my shoes, socks, pants with DEET. My scanning for butterflies as I walked along did not include my feet, but I could not help but notice when a bright red and blue butterfly the size of a healthy Mourning Cloak circled me suspiciously a couple times and then landed on my person. DEET is supposed to discourage insects, but this Agrias could not get enough. No doubt there were some electrolytes on my sweaty ankles, but electrolytes were saturating my shirt, too, so I can only conclude it was the DEET he was after. Eventually I got too close with my camera and it skipped off to perch head-down on a nearby tree trunk.Dicta Silverdrop (Epargyreus dicta: Hesperiidae: Eudaminae) Pantiacolla October 15, 2024 (photo by Steve Cary). New Mexico’s familiar Silver-spotted Skipper and Huachuca Skipper have many relatives in South America.Common Morpho (Morpho helenor: Nymphalidae: Morphinae) Pantiacolla; October 15, 2024 (photo by Steve Cary). Yes, Morphos were seen regularly. This palm-sized beauty is best known for its iridescent blue dorsum, which is very hard to catch well on film. Thankfully the ventrum is no less amazing.
Euselasia attrita or similar (Riodinidae: Riodininae) Pantiacolla; October 15, 2024 (photo by Steve Cary). Several related metalmarks have this same basic color pattern, even the hint of indigo peeking out from the dorsal forewing. E. attrita seemed like the best fit on hindwing marks, but other species cannot be ruled out. A collected specimen would give a full view of the dorsum and be easier to ID.
no English nameEuselasia orba spectralis Riodinidae: Riodininae) Pantiacolla; October 15, 2024 (photo by Steve Cary). Many metalmarks prefer to rest beneath leaves. This adds to the challenge of finding them and photographing them.
Vermiculated Leafwing or Glauce Leafwing (Memphis g. glauce Nymphalidae: Charaxinae) Pantiacolla; October 16, 2024 (photo by Steve Cary). There are dozens of Memphis species in Peru and the Amazon. Their undersides are amazing and variable dead leaf mimics. Uppersides tend to be the most diagnostic, but getting those photos takes luck and patience. They are closely related to our leafwings (Anaea species).
Trebula Groundstreak (Calycopis trebula: Lycaenidae: Theclinae) Pantiacolla; October 17, 2024 (photo by Steve Cary). Some butterflies are (understandably) flash-wary. I rarely use a flash in New Mexico, but in the cloudforest it is a necessity, so I did my best, deploying a fill flash on almost every shot. Some butterflies seem not to notice the flash, but others react, sometimes instantaneously, or so it seems. The flash has some duration, admittedly brief, but occasionally a butterfly’s quick reflex, results in a quick pic of a bug departing the scene as fast as it can. In this case the butterfly tipped over head-first. It seemed a bit humorous to me, but was undoubtedly a fright for the ‘streak, or at least an embarrassment :^)Amazonian Rose-Phantom or Blushing Phantom (Cithaerias aurorina Nymphalidae: Satyrinae) Pantiacolla; October 18, 2024 (photo by Steve Cary). It’s quite a special experience to watch the various phantom-satyrs flit patiently along the shady forest floor and trails . . .Emerald-Patched Cattleheart (Parides sesostris: Papilionidae: Papilioninae) Pantiacolla; October 18, 2024 (photo by Steve Cary). One of few swallowtail species seen on the entire trip.
We departed Pantiacolla on October 19, cruising back upstream on the boat that brought us: about 2.5 hours to travel ~15 miles. After docking back at the little port, we had a few minutes waiting for our van to come and get us. What did we do for those 30 minutes? We photographed butterflies of course. Once back in the van it was back to Manu Paradise Lodge, where we had stayed for the first several days of the trip.
Turquoise Emperor (Doxocopa laurentia Nymphalidae: Apaturinae) Manu Paradise Lodge; October 19, 2024 (photo by Steve Cary) you can’t make these up! related to our Emperors . . .Sylphina Angel (Chorinea sylphina Riodinidae: Riodininae) Morroleguia; October 20, 2024 (photo by Steve Cary). On our first full day back from the Pantiacolla “lowlands” we steered back up to some high country, stopping at a stream crossing at about 7000 feet elevation.Pink-Banded Longwing (Heliconius telesiphe: Nymphalidae: Heliconiinae) Morroleguia; October 20, 2024 (photo by Steve Cary).
We were due to head back to Cusco and home on 10/22, so of course the only way to spend 10/21 was at Quitacalzon.
Croniades auraria(Hesperiidae: Pyrrhopyginae) Quitacalzon; October 21, 2024 (photo by Steve Cary). Firetip Skippers like this one are an exciting part of the butterfly experience in this part of the world; I photographed 12 species on this trip. New Mexico has one species in this group, the Dull Firetip (Apyrrothrix araxes), flying only in the southernmost part of our Bootheel. Tawny Leafwing (Zaretis isidora Nymphalidae: Charaxinae) Quitacalzon; October 21, 2024 (photo by Steve Cary). Not a bad dead-leaf mimic.Acontius Greatstreak (Denivia acontius: Lycaenidae: Theclinae) Quebrada Quitacalzon; October 21, 2024 (photo by Steve Cary). This was fun because when I first took the photos I thought was a routine greatstreak like Denivia phegeus, which is regularly seen in the study area. I would have been happy to see that one, too, but Mike McInnis right away caught the white smudge toward the apex of the forewing and concluded it was this bug – a new species for the study area!no English name found (Cariomothis erotylus Riodinidae: Riodininae) Quitacalzon; October 21, 2024 (photo by Steve Cary). a very distinctive metalmark only recorded one time in the study area prior to our trip in October 2024.Hassan Greenstreak (Chalybs hassan Lycaenidae: Theclinae) Quebrada Quitacalzon; October 21, 2024 (photo by Steve Cary). The green hairstreaks do have a particular allure. This was a lifer for me. Taken on our last day at this fantastic site, it was one of several tiny hairstreaks that I captured “on film.” A good day indeed.Lathy’s Hairstreak (Lamasina lathyi Lycaenidae: Theclinae) Quitacalzon; October 21, 2024 (photo by Steve Cary). Named to honor our fearless leader, Gerardo Lamas.The Achilles, Taper-Lined, or Blue-Banded Morpho (Morpho achilles phokylides Nymphalidae: Morphiniae) Quitacalzon; October 21, 2024 (photo by Steve Cary). For the duration of this expedition, I kept my eyes open for Morphos and studied their behavior to figure how to best photograph their glorious blue uppersides. Underside photos are not hard to get because they perch with wings closed. Photographing one in flight is out of the question because they move too fast and too jerkily. When perched, they sometimes open their wings briefly. On a couple of occasions I put the camera on “burst mode” and just clicked off about 2 shots per second, waiting for the perched Morpho to open. That approach did not always work, but it did here. I was in its space and it was not pleased with me; it moved around a bit, seemingly agitated, Finally on exposure 102 it opened.
I think that is a representative cross-section of the butterflies I saw and some of the memorable experiences I had with them. I want to thank the other team members for their friendship and collegiality. I learned a lot from them and I hope to put that to use on a future trip to the Amazon!
7 thoughts on “Peruvian Butterflies: My October 2024 Snapshot”
Rebecca Gracey
Steve, What beautiful butterflies! Your photographs are a treat to view.
Thank you Steve. Not only for the wonderful photos and stories but for the memories of my past trips to the area – to video birds – that your blog brought to mind. I was always amazed at the beauty which would show up when I collapsed by the trail after carrying tripods and cameras with big lens all day. My theory was – “want to see butterflies?” – walk for hours through the jungle and fall down in a sweaty heap, in moments you will be covered with a beautiful array of rather passive butterflies.
Nice photos Steve, I knew right away those photos were not from your Inyo County, California trip and I am grateful that you did not ask me to identify any of these unfamiliar butterflies. And I am glad Bob Barnes is still around. He was a regular in our Kern-Tulare County, California butterfly counts, before those were discontinued
Hey Steve, thanks for posting these great images! As a member of the team I was unable to go on this trip so getting to see familiar “faces” is really nice. That new skipper is whacked out cool! Every day in the field down there is exciting knowing that you may stumble upon something “new”! It happened to me on the last trip not far from Paradise Lodge. I do not know if Olaf has published it yet but it was yet another new skipper.
Steve, What beautiful butterflies! Your photographs are a treat to view.
Amazing photos and descriptions. Your followers have been patiently waiting.
Thank you Steve. Not only for the wonderful photos and stories but for the memories of my past trips to the area – to video birds – that your blog brought to mind. I was always amazed at the beauty which would show up when I collapsed by the trail after carrying tripods and cameras with big lens all day. My theory was – “want to see butterflies?” – walk for hours through the jungle and fall down in a sweaty heap, in moments you will be covered with a beautiful array of rather passive butterflies.
Nice photos Steve, I knew right away those photos were not from your Inyo County, California trip and I am grateful that you did not ask me to identify any of these unfamiliar butterflies. And I am glad Bob Barnes is still around. He was a regular in our Kern-Tulare County, California butterfly counts, before those were discontinued
Thank you for sharing, Steve! What glorious little beauties you got to see and photograph! What a wonderful experience to treasure. ❤️
Hey Steve, thanks for posting these great images! As a member of the team I was unable to go on this trip so getting to see familiar “faces” is really nice. That new skipper is whacked out cool! Every day in the field down there is exciting knowing that you may stumble upon something “new”! It happened to me on the last trip not far from Paradise Lodge. I do not know if Olaf has published it yet but it was yet another new skipper.
Fantastic to read and look at your pictures! Thanks so much for posting this.